The Potato Virus

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Introduction
Potato virus X (PVX) is a plant RNA virus. Its infection cycle includes invasion of the host plant, RNA replication, translation of viral proteins, cell-to-cell movement and release of new virions. Upon infection, PVX releases its plus-strand RNA genome from the virion and produces a viral replicase using host translation machinery. The replicase synthesizes minus-strand RNA. Subsequently new plus-strand genomic RNA (gRNA) and subgenomic RNAs (sgRNAs) are produced. Movement and coat proteins are translated from the sgRNAs for encapsidation and movement through plasmodesmata. In this mini-review, all aspects of the virus infection cycle of PVX are discussed.

Classification
Potato virus X (PVX) is a small non-enveloped virus, which has a monopartite plus-strand RNA genome. Therefore it belongs to class IV of the Baltimore system. PVX is a member of the Potexvirus genus. Along with eight other genera, this genus belongs to the Alphaflexiviridae family [1].

Virion morphology and content
PVX forms small filamentous virions which have a deeply grooved surface [2]. The flexuous filaments consist of a single plus-strand genomic RNA coated with viral coat protein (CP). The helical structure has a pitch of 3.6nm and contains 8 7/8 CP subunits per turn [3]. The full-length filaments have a model length of 515nm with a diameter of 13nm [4].

CP consists of a single domain with high α-helical content [4]. The N-terminal part this domain is surface exposed whereas the C-terminal region buried in the virion. Several experiments indicate the CP is an O-glycoprotein. Equal amounts of galactose and fructose residues are O-linked to an acetylated serine residue at the N-terminal region [2]. This mediates the formation of a structured...

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...ut 10%. This is despite of obtaining high virus titers. Symptomless plants are regarded as carriers of PVX and are an important source of infection. Under more severe conditions, necrotic streaks and severe mosaic, crinkling and rugosity of the leaves can occur. Especially upon mixed infections with potato virus Y (PVY), the yield loss can be up to 50%.

Several potato plants carry resistance genes against PVX. Up to date two resistance genes have been identified. One gene product mediates salicylic acid systemic defense responses, whereas the other encodes the Rx protein. This protein has a nucleotide binding-site (NBS) and a leucine-rich repeat (LRR). The native fold of Rx confers an inactive conformation. Upon interactions with the viral CP, Rx partially unfolds, thereby its NBS-LRR domain [27]. This activates the signaling cascade mediating virus resistance.

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